Campanian
System/ Period | Series/ Epoch | Stage/ Age | Age (Ma) | |
---|---|---|---|---|
Paleogene | Paleocene | Danian | younger | |
Cretaceous | Upper/ Late | Maastrichtian | 66.0 | 72.1 |
Campanian | 72.1 | 83.6 | ||
Santonian | 83.6 | 86.3 | ||
Coniacian | 86.3 | 89.8 | ||
Turonian | 89.8 | 93.9 | ||
Cenomanian | 93.9 | 100.5 | ||
Lower/ Early | Albian | 100.5 | ~113.0 | |
Aptian | ~113.0 | ~125.0 | ||
Barremian | ~125.0 | ~129.4 | ||
Hauterivian | ~129.4 | ~132.9 | ||
Valanginian | ~132.9 | ~139.8 | ||
Berriasian | ~139.8 | ~145.0 | ||
Jurassic | Upper/ Late | Tithonian | older | |
Subdivision of the Cretaceous system according to the ICS, as of 2017.[1] |
The Campanian is, in the ICS' geologic timescale, the fifth of six ages of the Late Cretaceous epoch (or, in chronostratigraphy: the fifth of six stages in the Upper Cretaceous series). The Campanian spans the time from 83.6 ± 0.7 Ma to 72.1 ± 0.6 Ma (million years ago). It is preceded by the Santonian and it is followed by the Maastrichtian.[2]
The Campanian was an age when a worldwide sea level rise drowned many coastal areas. The morphology of some of these areas has been preserved as an unconformity beneath a cover of marine sedimentary rocks.[3][4]
Contents
1 Stratigraphic definition
1.1 Subdivision
2 Paleontology
2.1 †Ankylosaurs
2.2 Birds (avian theropods)
2.3 Bony fish
2.4 Cartilaginous fish
2.5 †Ceratopsians
2.6 Crocodylomorphs
2.7 Mammals
2.8 †Ornithopods
2.9 †Pachycephalosaurs
2.10 †Plesiosaurs
2.11 †Pterosaurs
2.12 †Sauropods
2.13 Squamates
2.14 Testudines
2.15 †Theropods (non-avian)
3 References
4 External links
Stratigraphic definition
The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the département Charente-Maritime. The original type locality was an outcrop near the village of Aubeterre-sur-Dronne in the same region. Due to changes of the stratigraphic definitions, this section is now part of the Maastrichtian stage.
The base of the Campanian stage is laid at the extinction of crinoid species Marsupites testudinarius. A GSSP had not yet been ratified in 2009. One possible candidate is in a section near a dam at Waxahachie, Texas.
The top of the Campanian is defined as the place in the stratigraphic column where the ammonite Pachydiscus neubergicus first appears.
Subdivision
The Campanian is sometimes subdivided into Lower, Middle and Upper subages. In the Tethys domain, the Campanian encompasses six ammonite biozones. They are, from young to old:
- zone of Nostoceras hyatti
- zone of Didymoceras chayennense
- zone of Bostrychoceras polyplocum
- zone of Hoplitoplacenticeras marroti/Hoplitoplacenticeras vari
- zone of Delawarella delawarensis
- zone of Placenticeras bidorsatum
Paleontology
During the Campanian age, a radiation among dinosaur species occurred. In North America, for example, the number of known dinosaur genera rises from 4 at the base of the Campanian to 48 in the upper part. This development is sometimes referred to as the "Campanian Explosion". However, it is not yet clear if the event is artificial, i.e. the low number of genera in the lower Campanian can be caused by a lower preservation chance for fossils in deposits of that age. The generally warm climates and large continental area covered in shallow sea during the Campanian probably favoured the dinosaurs. In the following Maastrichtian stage, the number of North American dinosaur genera found is 30% less than in the upper Campanian.[5]
Animals that lived in the Campanian include:
†Ankylosaurs
Ankylosaurs of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Aletopelta | Point Loma Formation, California, USA | A medium-sized ankylosaurid, estimated to be around 6 m (20 ft) long. | ||
Antarctopelta | Santa Marta Formation, James Ross Island, Antarctica | A stocky ankylosaur protected by armor plates embedded in the skin. Although a complete skeleton has not been found, the species is estimated to have reached a maximum length of 4 meters (13 feet). Displays characteristics of both ankylosaurids and nodosaurids. | ||
Edmontonia | Campanian to Maastrichtian | Horseshoe Canyon Formation, Alberta, Canada | A bulky nodosaurid at roughly 6.6 m (22 ft) long. It had small, ridged bony plates on its back and many sharp spikes along its body sides. The four largest spikes jutted out from the shoulders on each side, two of which were split into subspines in some specimens. Its skull had a pear-like shape when viewed from above. | |
Euoplocephalus | ||||
Nodocephalosaurus | ||||
Palaeoscincus | Judith River Formation | known from a single tooth | ||
Panoplosaurus | Judith River Formation, Alberta, Canada; Montana, USA | A 5.5–7 m long nodosaurid. | ||
Pinacosaurus | ||||
Saichania | ||||
Shanxia | ||||
Struthiosaurus | ||||
Tarchia | ||||
Tianzhenosaurus |
Birds (avian theropods)
Birds of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Hesperornis | ||||
Ichthyornis | ||||
Neogaeornis wetzeli | A marine bird from Chile. It had the midfeet of a foot-propelled diving bird, but its relationships are enigmatic. The only known species is from the Campanian-Maastrichtian boundary. |
Bony fish
Bony fish of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Xiphactinus |
Cartilaginous fish
Cartilaginous fish of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Chlamydoselachus | ||||
†Schizorhiza |
†Ceratopsians
Ceratopsians of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Achelousaurus
| 74.2 million years ago | |||
Agujaceratops
| 77 million years ago | |||
Albertaceratops
| ||||
Anchiceratops
| ||||
Avaceratops
| ||||
Bagaceratops
| ||||
Bainoceratops
| ||||
Brachyceratops
| ||||
Breviceratops
| ||||
Centrosaurus
| ||||
Cerasinops
| ||||
Ceratops
| ||||
Chasmosaurus
| ||||
Coahuilaceratops
| ||||
Coronosaurus
| ||||
Diabloceratops
| ||||
?Dysganus
| ||||
Einiosaurus
| ||||
Eoceratops
| ||||
Graciliceratops
| ||||
Gryphoceratops
| ||||
Judiceratops
| ||||
Kosmoceratops
| ||||
Lamaceratops
| ||||
Magnirostris
| ||||
Medusaceratops
| ||||
Mercuriceratops
| ||||
Mojoceratops
| ||||
Monoclonius
| ||||
Nasutoceratops
| ||||
?Notoceratops
| Chubut Province, Argentina | A dubious genus of possible ceratopsian affinity | ||
Pachyrhinosaurus
| ||||
Pentaceratops
| ||||
Platyceratops
| ||||
Prenoceratops
| ||||
Protoceratops
| ||||
Rubeosaurus
| ||||
Spiclypeus
| ||||
Spinops
| ||||
Styracosaurus
| ||||
Titanoceratops
| ||||
Udanoceratops
| ||||
Unescoceratops
| ||||
Utahceratops
| ||||
Vagaceratops
| ||||
Xenoceratops
|
Crocodylomorphs
Crocodylomorphs of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
†Deinosuchus |
Mammals
Mammals of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Alphadon | ||||
Didelphodon | ||||
Kamptobaatar | ||||
Kennalestes | ||||
Kryptobaatar | ||||
Zalambdalestes |
†Ornithopods
Ornithopods of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Aralosaurus | 85.8 mya | Asia | Aralosaurus was about the size of an elephant. Although very little is known about Aralosaurus (only one near complete skull has been found); it was identified by a beak with nearly 1,000 small teeth in 30 rows. These teeth were used for breaking up plant matter by chewing, a feature common in herbivorous dinosaurs, but unusual for reptiles.The back of an Aralosaurus skull was wide, a feature suggestive of large jaw muscles used to power its chewing apparatus. | |
Brachylophosaurus | 76.5 mya | Montana, USA; Alberta, Canada | Brachylophosaurus was a typical hadrosaur which reached an adult length of 9 meters (30 feet). | |
Corythosaurus | 77-76.5 mya | Alberta, Canada | Corythosaurus weighed in at 4 tonnes and measured roughly 10 metres (33 feet) from nose to tail. Like other hadrosaurs it had a toothless beak, the back of the jaws contained a dental battery composed of hundreds of small, interlocking teeth. These were used to crush and grind plant matter and were continually replaced as they wore away. | |
Diclonius | 75 mya | Montana, USA | ||
Edmontosaurus | 73.0-76.5 mya | Canada | Edmontosaurus included some of the largest hadrosaurid species, measuring up to 12 metres (39 feet) long and weighing around 4.0 metric tons (4.4 short tons). | |
Gasparinisaura | 85 mya | Argentina | Gasparinisaura was a small bipedal herbivore. In 2010 Gregory S. Paul estimated the length at 1.7 metres, the weight at thirteen kilogrammes. | |
Gilmoreosaurus | 72 mya | Mongolia | ||
Gryposaurus | 83-75.5 mya | Alberta, Canada | Gryposaurus was a hadrosaurid of typical size and shape. | |
Hadrosaurus | 79.5 mya | New Jersey, USA | It was likely bipedal for the purposes of running, but could use its forelegs to support itself while feeding. | |
Hypacrosaurus | 75-67 mya | Alberta, Canada | Hypacrosaurus is most easily distinguished from other hollow-crested duckbills by its tall neural spines and the form of its crest. The neural spines, which project from the top of the vertebrae, are 5 to 7 times the height of the body of their respective vertebrae in the back,[4] which would have given it a tall back in profile. The skull's hollow crest is like that of Corythosaurus, but is more pointed along its top, not as tall, wider side to side, and has a small bony point at the rear | |
Hypsibema | North Carolina and Missouri, USA | |||
Kritosaurus | 73 mya | North America | The type specimen of Kritosaurus navajovius is only represented by a partial skull and lower jaws, and associated postcranial remains. | |
Lambeosaurus | 76-75 mya | Alberta, Canada | ||
Lophorhothon | 80 mya | Alabama, USA | ||
Maiasaura | 76.7 mya | Montana, USA | Maiasaura was large, attaining an adult length of about 9 metres (30 feet) and had the typical hadrosaurid flat beak and a thick nose. It had a small, spiky crest in front of its eyes. The crest may have been used in headbutting contests between males during the breeding season. | |
Mandschurosaurus | Asia | |||
Microhadrosaurus | China | |||
Mochlodon | Austria | A rhabdodontid. | ||
Naashoibitosaurus | 73 mya | New Mexico, USA | Naashoibitosaurus, based as it is on a single partial skeleton, is not well known in terms of anatomy. Its skull, the most thoroughly described portion, has a low nasal crest that peaks in front of the eyes, but does not strongly arch as in Gryposaurus. | |
Nipponosaurus | Russia | |||
Orodromeus | 76.7 mya | Montana, USA | Orodromeus was a small fast bipedal herbivore that probably coexisted with dinosaurs such as Daspletosaurus and Einiosaurus. Its length was estimated by Horner & Weishampel at 2.5 metres. | |
Parasaurolophus | 76.5-73 mya | Alberta, Canada; New Mexico and Utah, USA | ||
Prosaurolophus | 76-75 mya | Alberta, Canada | Prosaurolophus was a large-headed duckbill; the most complete described specimen has a skull around 0.9 meters (3.0 feet) long on a ~8.5 meter long skeleton (~28 ft).[2] It had a small, stout, triangular crest in front of the eyes; the sides of this crest were concave, forming depressions. The upper arm was relatively short. | |
Pteropelyx | Montana, USA | |||
Rhabdodon | 72 mya | France; Spain; Haţeg Island, Romania | It is unclear whether it was an iguanodont or a hypsilophodont, and may be a "missing link" between the two. Current evidence indicates it is an iguanodont similar to Tenontosaurus. | |
Saurolophus | 69.5-68.5 mya | North America, Asia | Saurolophus is known from material including nearly complete skeletons, giving researchers a clear picture of its bony anatomy. S. osborni, the rarer Albertan species, was around 9.8 meters (32 feet) long, with its skull a meter long (3.3 feet). Its weight is estimated at 1.9 tonnes (2.1 tons). S. angustirostris, the Mongolian species, was larger; the type skeleton is roughly 12 meters (39 feet) long, and larger remains are reported. | |
Shantungosaurus | 72 mya | China | It is one of the longest and largest known hadrosaurids; the composite skeleton of a medium-sized individual mounted at the Geological Institute of China in Beijing measures 14.72 metres (48.3 feet) in length. | |
Stephanosaurus | Dinosaur Park Formation, Alberta | |||
Tanius | China | |||
Trachodon | 77 mya | Montana, USA | ||
Tsintaosaurus | 72 mya | Southern China | ||
Velafrons | 72 mya | Mexico |
†Pachycephalosaurs
Pachycephalosaurs of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Alaskacephale | Prince Creek Formation, Alaska, USA | |||
Colepiocephale | Alberta, Canada | The oldest known pachycephalosaurid. | ||
Goyocephale | Mongolia | |||
Gravitholus | ||||
Hanssuesia | Alberta, Canada; Montana, USA | Distinguished from other pachycephalosaurs by having a depressed parietal region, wide frontoparietal dome, broad nasal characteristics, reduced prefontal lobes, and a reduced parietosquamosal shelf. | ||
?Heishansaurus | ||||
Homalocephale | Mongolia | Sporting a flat, wedge-shaped skull roof, Homalocephale was different from other pachycephalosaurs. | ||
?Micropachycephalosaurus | ||||
Ornatotholus | ||||
Prenocephale | ||||
Sphaerotholus | ||||
Stegoceras | ||||
Tylocephale | ||||
Wannanosaurus |
†Plesiosaurs
Plesiosaurs of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
†Elasmosaurus | 80.5 mya | Pierre Shale, Kansas, USA | Elasmosaurus is a genus of plesiosaur with an extremely long neck. | |
†Styxosaurus | 83.5-80.5 mya | Logan County, Kansas | Styxosaurus is an Elasmosaurid plesiosaur. |
†Pterosaurs
Pterosaurs of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Aerotitan | Campanian-Maastrichtian | Allen Formation, Patagonia, Argentina | ||
Bogolubovia | Rybushka Formation, Petrovsk, Russia | |||
Geosternbergia | USA, North America | Geosternbergia was originally a species of Pteranodon and is famous for its oddly shaped crest. | ||
Montanazhdarcho | Montana, USA | Small azhdarchoid pterosaur, probably a tapejarid | ||
Navajodactylus | New Mexico, USA, and Alberta, Canada | Known primarily from forearm elements; tentatively assigned to Azhdarchidae, though most likely not part of it. | ||
Nyctosaurus | mid-western United States | Nyctosaurus is a genus of pterodactyloid pterosaur. | ||
Piksi | Montana, USA | Piksi is a genus of pterosaurs containing the single species Piksi barbarulna. | ||
Pteranodon | Kansas, USA, North America | Pteranodon is a genus of pterosaurs which included some of the largest known flying reptiles, with wingspans over 6 metres | ||
Quetzalcoatlus | Texas, USA | Quetzalcoatlus was a pterodactyloid pterosaur known from the Late Cretaceous of North America and one of the largest known flying animals of all time. | ||
Volgadraco | Saratov, Russia | Azhdarchid pterosaur. |
†Sauropods
Sauropods of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Alamosaurus | Southwestern United States | Alamosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now North America. It was a large quadrupedal herbivore. | ||
Andesaurus | Neuquén Province, Argentina | Andesaurus is a genus of basal titanosaurian sauropod dinosaur. | ||
Dreadnoughtus | Cerro Fortaleza Formation, Argentina | Dreadnoughtus is one of the largest titanosaurs known. | ||
Gondwanatitan | Adamantina Formation and Cambabe Formation, Brazil | |||
Huabeisaurus | North East, China | A member of the Euhelopodidae sauropods. | ||
Laplatasaurus | Allen Formation and Anacleto Formation, both in Argentina; Palacio Formation, Uruguay | |||
Loricosaurus | Campanian-Maastrichtian | Allen Formation, Argentina | ||
Microcoelus | Santonian-Campanian | Bajo de la Carpa Formation, Argentina | ||
Neuquensaurus | Anacleto Formation, Argentina | |||
Overosaurus | Neuquén Province, Argentina | Small-sized titanosaur. | ||
Quaesitosaurus | Shar Tsav, Mongolia | Quaesitosaurus is a genus of titanosaurian sauropod. | ||
Saltasaurus | north-west Argentina; Uruguay | Saltasaurus is a genus of titanosaurid sauropod dinosaur. An estimated length of 12 metres (39 feet) and a mass of 7 tonnes (8 tons). | ||
Rocasaurus | Campanian-Maastrichtian | Allen Formation, Rio Negro Province, Argentina |
Squamates
Squamates of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Halisaurus | ||||
Mosasaurus | ||||
Plotosaurus | ||||
Taniwhasaurus | New Zealand, Japan, Antarctica | |||
Tylosaurus |
Testudines
Testudines of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
†Archelon |
†Theropods (non-avian)
David J. Varrichio observes that during the late Campanian Alberta and Montana had very similar theropods despite significant differences in the types of herbivorous dinosaur faunas.[6]
Non-avian theropods of the Campanian | ||||
---|---|---|---|---|
Taxa | Presence | Location | Description | Images |
Abelisaurus | Allen Formation?, Anacleto Formation?, Argentina | Bipedal carnivore that probably reached 7 to 9 meters in length; known from only one partial skull. | ||
Albertosaurus | ||||
Appalachiosaurus | ||||
Archaeornithomimus | ||||
Bambiraptor | ||||
Byronosaurus | ||||
Citipati | ||||
Carnotaurus | ||||
Chirostenotes | ||||
Daspletosaurus | ||||
Deinodon | Judith River Formation | |||
Dromaeosaurus | ||||
Dromiceiomimus | ||||
Dryptosaurus | ||||
Gobivenator | ||||
Gorgosaurus | ||||
Harpymimus | ||||
Khaan | ||||
Kol | ||||
Linheraptor | ||||
Lythronax | Wahweap Formation, Utah | A 7-meter tyrannosaurid known from a partially complete skull, some vertebrae and a complete pubis | ||
Luanchuanraptor | ||||
Mahakala | ||||
Nanshiungosaurus | ||||
Noasaurus | ||||
Ornithomimus | ||||
Oviraptor | Mongolia | |||
Parvicursor | ||||
Pyroraptor | Var, France | |||
Saurornithoides | ||||
Saurornitholestes | ||||
Shuvuuia | ||||
Struthiomimus | ||||
Troodon | ||||
Tsaagan | ||||
Variraptor | Var, France | |||
Velociraptor | Mongolia and China | |||
Xenotarsosaurus | ||||
Zhuchengtyrannus | Wangshi Group, Zhucheng, China | One of the largest tyrannosaurids at between 10–12 meters. Known from a lower jaw and maxilla slightly smaller than those of the later Tyrannosaurus. |
References
^ http://www.stratigraphy.org/index.php/ics-chart-timescale
^ See Gradstein et al. (2004) for a detailed version of the geological timescale
^ Lidmar-Bergström, Karna; Bonow, Johan M.; Japsen, Peter (2013). "Stratigraphic Landscape Analysis and geomorphological paradigms: Scandinavia as an example of Phanerozoic uplift and subsidence". Global and Planetary Change. 100: 153–171. doi:10.1016/j.gloplacha.2012.10.015..mw-parser-output cite.citation{font-style:inherit}.mw-parser-output .citation q{quotes:"""""""'""'"}.mw-parser-output .citation .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Lock-green.svg/9px-Lock-green.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .citation .cs1-lock-limited a,.mw-parser-output .citation .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Lock-gray-alt-2.svg/9px-Lock-gray-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .citation .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/a/aa/Lock-red-alt-2.svg/9px-Lock-red-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration{color:#555}.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration span{border-bottom:1px dotted;cursor:help}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Wikisource-logo.svg/12px-Wikisource-logo.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output code.cs1-code{color:inherit;background:inherit;border:inherit;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;font-size:100%}.mw-parser-output .cs1-visible-error{font-size:100%}.mw-parser-output .cs1-maint{display:none;color:#33aa33;margin-left:0.3em}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration,.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-right{padding-right:0.2em}
^ Surlyk, Finn; Sørensen, Anne Mehlin (2010). "An early Campanian rocky shore at Ivö Klack, southern Sweden". Cretaceous Research. 31: 567–576. doi:10.1016/j.cretres.2010.07.006.
^ See Weishampel et al. (2004)
^ "Abstract," in Varricchio (2001). Page 42.
.mw-parser-output .smallcaps{font-variant:small-caps}
Gradstein, F.M.; Ogg, J.G. & Smith, A.G.; 2004: A Geologic Time Scale 2004, Cambridge University Press.- Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana.
Weishampel, D.B.; Barrett, P.M.; Coria, R.A.; Le Loueff, J.; Xu, X.; Zhao, X.; Sahni, A.; Gomani, E.M.P. & Noto, C.N.; 2004: Dinosaur distribution, in:
Weishampel, D.B.; Dodson, P. & Osmólska, H. (eds.): The Dinosauria, University of California Press, Berkeley (2nd ed.),
ISBN 0-520-24209-2, pp 517–606.
External links
- GeoWhen Database - Campanian
Late Cretaceous timescale, at the website of the subcommission for stratigraphic information of the ICS
Stratigraphic chart of the Late Cretaceous, at the website of Norges Network of offshore records of geology and stratigraphy- Campanian Microfossils: 75+ images of Foraminifera